Hulthemia blotch eluding mendelian law classification

The blotch is eluding a simple classification by means of the mendelian laws.
The blotch maybe inside but not always visible.

R. persica has six layers of palisade parenchyma and I am wondering where the blotch is located in hybrid hulthemia.

Not at top in case of this open pollinated seedling of a hybrid hulthemia.

[attachment 1864 hybrid_hulthemia_a.jpg]
[attachment 1865 hybrid_hulthemia_b.jpg]
[attachment 1866 hybrid_hulthemia_c.jpg]

There is a light yellow tinted layer on top, fading to white.
Potentially this layer ends transparent.

Intercellular holes reflect white light until they collapse.
Finally the layer containing the blotch is visible.

In fact, statements about allele dominance of the blotch are quite unsure if authors did not recognize these circumstances.

See this link for information on
COMPARATIVE ANATOMEY OF LEAF
AND RACHIS OF ROSA L.
(ROSACEAE) IN IRAN AS TAXONOMICAL IMPLICATION

I would kindly ask you to read this document carefully and provide the forum members with your comments and suggestions.

Hello Eyecatcher,

Thank you for the article. It is very interesting. To me it shows that there are several physical characteristics on a more microscopic level that help to distinguish the various species from one another. I am uncertain though how this article relates to the Hulthemia blotch.

Regarding the question of the blotch, I agree that there is much to be still figured out. To me it appears that the blotch gene(s) interact with rose genes such that it seems to be a dominant characteristic with some crosses while perhaps recessive when other rose genes are present. Also, the degree of blotch expression varies considerable from seedling to seedling.

With regard to your photos, the red coloration at the base of the petals on the older bloom can sometimes appear in non-hulthemia seedlings as they age when the weather is cooler. When looking at the blotch pigment under lower handheld magnification, to me the blotch pigment often seems more superficial than some of the other petal pigments.

Hopefully there will be more research to answer these questions about blotch expression.

That “red fade blotch” seems to have originated from the combination of Baby Chateau, Crimson Glory and Charlotte Armstrong. Years ago, when I researched the Halo Roots article about Ralph Moore’s Halo roses, I searched for common ancestors for all which demonstrated the lavender petal bases the Halos were bred from. Those three are the only common ancestors to all of them. Which added what, I have no idea, but they all figure in the lavender petal bases which have been selected to become red. Jim, your First Impression also contains them and has passed on the red fade blotch to my First Impression X April Mooncrest crested seedling. It begins blush pink and may retain the blush when it’s cooler. Heat bleaches it to pure white, but as it ages, the red petal base blotch appears. [flickr_photo src=http://farm4.staticflickr.com/3669/9225653895_b893e5109a.jpg nsid=67995840@N04 id=9225653895]DSCN4823[/flickr_photo][flickr_photo src=http://farm3.staticflickr.com/2856/9110906752_a3f0e7284f.jpg nsid=67995840@N04 id=9110906752]DSCN4599[/flickr_photo][flickr_photo src=http://farm4.staticflickr.com/3813/9240466135_22b0be0555.jpg nsid=67995840@N04 id=9240466135]DSCN4851[/flickr_photo][flickr_photo src=http://farm3.staticflickr.com/2834/9243247894_d8d44a13e3.jpg nsid=67995840@N04 id=9243247894]DSCN4853[/flickr_photo][flickr_photo src=http://farm6.staticflickr.com/5518/9243247202_b6f42cf2bb.jpg nsid=67995840@N04 id=9243247202]DSCN4855[/flickr_photo][flickr_photo src=http://farm6.staticflickr.com/5510/9243247612_2e0a0f362d.jpg nsid=67995840@N04 id=9243247612]DSCN4854[/flickr_photo]